^pKey;
Laboratory;
of;
Systematic;
and;
Evolutionary;
Botany,;
Institute;
of;
Botany,;
Chinese;
Academy;
of;
Sciences,;
Beijing;
100093,;
China;
^pUniversity;
of;
the;
Chinese;
Academy;
of;
Sciences,;
Beijing;
100049,;
China;
^pof;
Biology,;
University;
College;
London,;
London;
WC;
1E;
6BT,;
United;
Kingdom;
^pfor;
Computational;
and;
Evolutionary;
Biology,;
Institute;
of;
Zoology,;
Chinese;
Academy;
of;
Sciences,;
Beijing;
100101,;
China;