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Miocene abyssochrysoid gastropod Provanna from Japanese seep and whale-fall sites

机译:来自日本渗漏和鲸鱼坠落地点的中新世abyssochrysoid腹足类provanna

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摘要

We describe three Miocene species of Provanna from Japan, two new and one in open nomenclature, that represent the only known fossil examples from whale-falls and a considerable increase in the Miocene diversity of the genus. Provanna hirokoae sp. nov. comes from the latest Middle Miocene Kuroiwa seep site in central Honshu. The shells of this species are mostly recrystallized, but contain relict crossed lamellar microstructures. Provanna alexi sp. nov. is from the early Middle Miocene Shosanbetsu whale-fall site in northwestern Hokkaido, and has well preserved shells comprising an outer simple prismatic layer and an inner crossed lamellar layer. The two Provanna specimens from the Middle Miocene Rekifune whale-fall site, in eastern Hokkaido, are preserved as external moulds only, so are left in open nomenclature. Based on current knowledge, the presence of an outer prismatic layer and an underlying crossed lamellar layer seems to be a common feature in the shells of Provanna, as well as in other genera belonging to the family Provannidae and the superfamily Abyssochrysoidea. Although the oldest occurrence of Provanna was in the Late Cretaceous, the genus did not spread geographically and ecologically until the Miocene (with four, or possibly five species), a date concordant with some molecular estimates. However, this could be an artefact of the fossil record because the known pre-Miocene seep and whale-falls are more geographically restricted than those from the Miocene.
机译:我们描述了来自日本的普罗旺纳地区的三种中新世物种,其中两种是新物种,一种是开放命名法,它们代表了鲸鱼降落和该属中新世多样性显着增加的唯一已知化石实例。 Provanna hirokoae sp。十一月来自本州中部最新的中新世黑岩岩渗流场。该种类的壳大部分是重结晶的,但含有残留的交叉的层状微结构。 Provanna alexi sp。十一月来自北海道西北部的中新世初山松山别鲸降落地点,壳保存完好,壳包括一个外部简单的棱柱层和一个内部交叉的层状层。来自北海道东部中新世中期风鲸降落地点的两个普罗旺那标本仅保存为外部霉菌,因此保留为开放式命名法。根据目前的知识,在Provanna以及其他属于Provannidae家族和Abyssochrysoidea家族的其他属中,外棱柱层和下面的交叉层状层的存在似乎是它们的共同特征。尽管普罗旺纳最古老的发生在白垩纪晚期,但是该属直到中新世(有四个或可能五个物种)才在地理和生态上传播,这一日期与某些分子估计相符。但是,这可能是化石记录的人工产物,因为与中新世相比,已知的中新世之前的渗流和鲸鱼落在地理上的限制更大。

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    Amano K; Little CTS;

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  • 年度 2014
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