^pLaboratory;
of;
Adaptation;
and;
Evolution;
of;
Plateau;
Biota,;
Northwest;
Institute;
of;
Plateau;
Biology,;
Chinese;
Academy;
of;
Sciences,;
Xining;
810008,;
China;
^pof;
Ecology,;
and;
Key;
Laboratory;
for;
Earth;
Surface;
Processes;
of;
Ministry;
of;
Education,;
Peking;
University,;
Beijing;
100871,;
China;