Key;
Laboratory;
of;
Genetic;
Resources;
and;
Evolution,;
Kunming;
Institute;
of;
Zoology,;
Chinese;
Academy;
of;
Sciences,;
Kunming;
650223;
China;
and;
Mathematics;
College,;
Yunnan;
University;
of;
Finance;
and;
Economics,;
Kunming;
650221,;
China;
of;
Zoology;
and;
Entomology,;
Rhodes;
University,;
Grahamstown;
6140,;
South;
Africa;
of;
Biology,;
University;
of;
Leeds,;
Leeds,;
LS2;
9JT,;
UK;