College of Eco-Environmental Engineering Guizhou Minzu University Guiyang 550025 China;
College of Eco-Environmental Engineering Guizhou Minzu University Guiyang 550025 China;
College of Eco-Environmental Engineering Guizhou Minzu University Guiyang 550025 China;
Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations Kunming Institute of Botany Kunming 650201 China;
The College of Humanities and Science of Guizhou Minzu University Guiyang 550025 China;