This paper is concerned with the bifurcation of limit cycles in general quadratic perturbations of quadratic codimension-four centres Q_4. Gavrilov and Iliev set an upper bound of eight for the number of limit cycles produced from the period annuli around the centre. Based on Gavrilov-Iliev's proof, we prove in this paper that the perturbed system has at most five limit cycles which emerge from the period annuli around the centre. We also show that there exists a perturbed system with three limit cycles produced by the period annuli of Q_4.
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