Analysing structural diversity in two temperate forests in northeastern China. (Special Issue: Forest observational studies: 'Data sources for analysing forest structure and dynamics'.)

摘要

This contribution presents an analysis of the structural diversity in two temperate forest ecosystems in northeastern China, based on two large contiguous observational studies covering an area of 5.2 ha (260x200 m2) each. The total number of living trees in the conifer and broad-leaved forest (CBF) study area is 2927 per ha, comprising 37 species and 21 genera. The old growth forest (OGF) study area has 2276 trees per ha, including 22 species and 13 genera. Tree species are classified according to their community status as mature and immature canopy, subcanopy and shrub. A clustering process based on two distinct communities of the bivariate dbh/height distributions is used to differentiate between mature and immature canopy species. Numerical analysis is based on these four distinct cohorts. Despite advances in remote sensing, mapped tree data in large observation windows are very rare. Thus, we are able to use methods for analysing forest structure which are suitable for both, unmapped and mapped data. The two data sets are unique in that all (approximately) 27,000 tree heights are available. Accordingly, it was possible to fit bimodal height distributions and bivariate mixed dbh/height distributions to almost all individual species that were represented by sufficiently large numbers. Methods of second order statistics (SOC), including marked point statistics as well as nearest neighbor statistics (NNS) based on nearest neighbourhood structure units are also presented, including bivariate mixtures of the NNS attributes "mingling" and "dominance". Mark correlations were investigated for several marks, including diameters, heights and nearest neighbor statistics. Finally, we discuss the most important results and motivate the need for detailed assessments in large contiguous field plots. The literature on spatial statistics is often rather technical, and there is relatively little exchange between mathematicians developing the theory and ecologists who have interesting data from observational studies, such as presented in this contribution.