Revised Classification of the Species within the Dryopteris variaComplex (Dryopteridaceae) in Japan

摘要

The Dryopteris varia complex (subg. Erythrovariae sect. Variae Fraser-Jenk.) is an apogamous ferncomplex. Most species of the complex are triploid apogamous, show morphological and genetic variation, and present difficulties in their identification. Recent molecular research suggested that the reasonthat each apogamous species in the complex is so variable is due to reticulate evolution through hybridization between diploid sexual and triploid apogamous species. In this study, we summarized the taxonomic treatment of 11 species within the D. varia complex (D. bissetiana, D. chichisimensis, D. erythrovaria sp. nov., D. hikonensis, D. insularis, D. kobayashii, D. protobissetiana, D. sacrosancta, D. saxifraga, D. subhikonensis sp. nov., and D. varia) on the basis of their genome constituents revealed bymolecular analyses.