New Albian macro- and palynoflora from the Negev (Israel) with description of a new gymnosperm morphotaxon

摘要

New plant macrofossil localities are found in the middle-upper part of the Albian Upper Hatira Formation of Makhtesh Qatan, an erosion crater in the northern Negev. These are so far the only outcrop localities in the Negev containing plant compressions and well-preserved sporomorphs. Their age assignments are controlled by the ammonite Knemiceras records below and above the plant-bearing sequence in the adjacent Makhtesh Hatira. The macrofossil assemblages are strikingly different in the sandy fluvial and shaly lacustrine facies, the former being dominated by Araucaria, Athrotaxopsis, Brachyphyllum and pinnatifid Sapindopsis, comparable at the plant assemblage and leaf morphological levels to the early to early late Albian assemblage of the North American Potomac flora. The lacustrine shale macrofossil assemblages appear archaic on account of their diverse pteridophyte component and sparse angiosperm remains. A new supposedly gnetophytic genus and species Qataniaria noae Krassilov, gen. et sp. nov. is dominant in the shale horizon. The sporomorph assemblages are strongly dominated by psilate trilete spores (87-93%). In the absence of elaterate forms, the angiosperm pollen Afropollis jardinus, Pennipollis, Tricolpites spp. and Walkeripollis sp. is consistent with the early Albian age. The abundance and diversity of conifers and the prominence of the fern bog assemblage suggest a relatively humid phase of the regional Albian climates.