In 1998, the second author of this paper raised the problem of classifying the irreducible characters of S_n of prime power degree. Zalesskii proposed the analogous problem for quasi-simple groups, and he has, in joint work with Malle, made substantial progress on this latter problem. With the exception of the alternating groups and their double covers, their work provides a complete solution. In this article we first classify all the irreducible characters of S_n of prime power degree (Theorem 2.4), and then we deduce the corresponding classification for the alternating groups (Theorem 5.1), thus providing the answer for one of the two remaining families in Zalesskii's problem. This classification has another application in group theory. With it, we are able to answer, for alternating groups, a question of Huppert: which simple groups G have the property that there is a prime p for which G has an irreducible character of p-power degree >1 and all of the irreducible characters of G have degrees that are relatively prime to p or are powers of p?
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