In this article, I present two conceptual problems for Craver's mutual manipulability account of constitutive relevance in mechanisms. First, constitutive relevance threatens to imply causal relevance despite Craver (and Bechtel)'s claim that they are strictly distinct. Second, if (as is intuitively appealing) parthood is defined in terms of spatio-temporal inclusion, then the mutual manipulability account is prone to counterexamples, asIshowby a caseofendosymbiosis.Ialso present a methodological problem (a case of experimental underdetermination) and formulate two partial, but fallible solutions based on the notions of parthood and synchronicity.
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