Shanghai Chenshan Plant Science Research Center Chinese Academy of Sciences Shanghai Chenshan Botanical Garden Songjiang Shanghai 201602 China;
School of Life Sciences East China Normal University Minhang Shanghai 200241 China;
Harvard University Herbaria 22 Divinity Avenue Cambridge MA 02138 USA;
Shanghai Chenshan Plant Science Research Center Chinese Academy of Sciences Shanghai Chenshan Botanical Garden Songjiang Shanghai 201602 China;