Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany Chinese Academy of Science Kunming 650201 China;
Plant Germplasm and Genomics Center The Germplasm Bank of Wild Species Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China;
Institute of Tibetan Plateau Research at Kunming Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China;
University of the Chinese Academy of Sciences Beijing 100049 China;