National Forest Ecosystem Observation & Research Station of Nyingchi Tibet Institute of Plateau Ecology Tibet Agriculture & Animal Husbandry University Linzhi 860000 China;
Key Laboratory of Forest Ecology in Tibet Plateau Tibet Agriculture & Animal Husbandry University Linzhi 860000 China;
Schoolof Life Sciences Lanzhou University Lanzhou 730000 China;
Schoolof Life Sciences Lanzhou University Lanzhou 730000 China;
Schoolof Life Sciences Lanzhou University Lanzhou 730000 China;
Schoolof Life Sciences Lanzhou University Lanzhou 730000 China;
Schoolof Life Sciences Lanzhou University Lanzhou 730000 China;
Tibet Plateau Institute of Biology Lhasa 850000 China;
Schoolof Life Sciences Lanzhou University Lanzhou 730000 China;