A conveyor (20) comprising at least one belt (22) defining an outer conveying surface divided into at least two laterally offset regions in which the rollers (50) in one region rotate in a first direction and the rollers (51) in a second region rotate in a second direction as the conveyor belt (22) advances in a conveying direction. The rollers (50, 51) rotate by rolling along roller-engagement surfaces (54) underlying the conveyor belt along a carryway. The rollers rotating in the first direction rotate with a velocity component in the conveying direction (32) different from the velocity component in the conveying direction of the rollers rotating in the second direction. An article positioned simultaneously atop rollers in both regions is rotated by the rotation of the rollers as the conveyor belt advances.
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