Revision of the earthworm genus Archipheretima Michaelsen(Clitellata: Megascolecidae), with descriptions of new speciesfrom Luzon and Catanduanes Islands, Philippines

摘要

A revision of Archipheretima (Clitellata: Megascolecidae) based on all available type material and the description of fivenew species modifies the genus diagnosis in the following points: Archipheretima have calciferous lamellar regions of theesophagus in the region of segments xi–xiii, paired supraesophageal vessels with connectives to the calciferous lamellae,hearts of xiii attached post-septally, multiple paired dorsolateral intestinal caeca, and usually very sparse dorsal setae. Bluecoloration is common to all the species described, and probably to the other known species—Von-Philippine speciespreviously assigned to the genus are excluded by this diagnosis; as a result, the new combinations Polypheretima picta(Michaelsen, 1892) and P. beccarii (Cognetti, 1909) are proposed. The redefined Archipheretima is a biogeographically andmorphologically homogenous taxon. The new species are named Archipheretima gritzae, A. middletoni, A. ricei, A. cofini,and A. pandanophila. They were collected only on Luzon Island, except A. gritzae which also occurs on Catanduanes. Thefull results, illustrated species descriptions, and discussions are given in the online edition of this paper.