A graph drawing is called a generalized thrackle if every pair of edges meets an odd number of times. In a previous paper, we showed that a bipartite graph G can be drawn as a generalized thrackle on an oriented closed surface M if and only if G can be embedded in M. In this paper, we use Lins' notion of a parity embedding and show that a non-bipartite graph can be drawn as a generalized thrackle on an oriented closed surface M if and only if there is a parity embedding of G in a closed non-orientable surface of Euler characteristic (X)( M) - 1. As a corollary, we prove a sharp upper bound for the number of edges of a simple generalized thrackle.
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