Reply to the Paper 'Beutel et al. 2018. Is dagger Skleroptera (dagger Stephanastus) an order in the stemgroup of Coleopterida (Insecta)?'

摘要

The recent critical revision of the order Skleroptera by Beutel et al. (2018a) is considered. We show several defects in their interpretation, contradicted by the original descriptions and diagnosis (Nel et al., 2013; Kirejtshuk & Nel, 2013). The main arguments of the initial interpretation of Stephanastus polinae Kirejtshuk et Nel, 2013 (Stephanastidae) and reasons for the proposal of the order Skleroptera (Kirejtshuk & Nel, 2013) are mentioned, together with reasons to reject the proposal of Beutel et al. (2018a, b). A comparison of Skleroptera with other neopteran orders is made. Lastly the problem of the position of Umenocoleus as a member of Coleopterida versus a roachoid is discussed, the first hypothesis being supported by synapomorphies which is not the case for the second.