Oryctoantiquus borealis, new genus and species from the eocene of Oregon, USA, the world's oldest fossil dynastine and largest fossil Scarabaeid (Coleoptera : Scarabaeidae : Dynastinae)

摘要

Oryctoantiquus borealis Ratcliffe and Smith is described as a new genus and new species of fossil dynastine Scarabaeidae from West Branch Creek of the Middle Eocene Clarno Formation (44.6-46.8 Ma) of north-central Oregon, U.S.A. This specimen is the oldest specimen attributable to the subfamily Dynastinae and the largest fossil in the family Scarabaeidae.