Author contributions: J.A.T. and D.S.B. designed research; J.A.T., J.D.R., R.A.K., T.T.M., C.J.B., and D.S.B. performed research; J.A.T., R.A.K., and D.S.B. analyzed data; J.A.T. and D.S.B. wrote the first draft of the paper; J.A.T., J.D.R., R.A.K., C.J.B., F.E.D., and D.S.B. edited the paper; D.S.B. wrote the paper.
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