State Key Laboratory of Systematic and Evolutionary Botany Institute of Botany Chinese Academy of Sciences Beijing 100093 China;
University of Chinese Academy of Sciences Beijing 100049 China;
State Key Laboratory of Systematic and Evolutionary Botany Institute of Botany Chinese Academy of Sciences Beijing 100093 China;
Shenzhen Key Laboratory for Orchid Conservation and Utilization The National Orchid Conservation Center of China and The Orchid Conservation and Research Center of Shenzhen Shenzhen 518114 China;
Key Laboratory of National Forestry and Grassland Administration for Orchid Conservation and Utilization at the College of Landscape Architecture Fujian Agriculture and Forestry University Fuzhou 350002 China;
College of Forestry and Landscape Architecture South China Agricultural University Guangzhou 510640 China;
Fujian Colleges and Universities and Engineering Research Institute of Conservation and Utilization of Natural Bioresources College of Forestry Fujian Agriculture and Forestry University Fuzhou 350002 China;
State Key Laboratory of Systematic and Evolutionary Botany Institute of Botany Chinese Academy of Sciences Beijing 100093 China;
Sino-Africa Joint Research Center Chinese Academy of Sciences Wuhan 430074 China;
