Key Laboratory of Biology and Genetic Resources of Tropical Crops of Tropical Bioscience and Biotechnology Academy of Tropical Agricultural Sciences 571101.R.China;
Key Laboratory for Biology and Genetic Resources of Tropical Crops of Hainan Province Institute for Tropical Agricultural Resources 571101.R.China;
Cash Crops Research Institute Academy of Agricultural Sciences 530007.R.China;
College of Tropical Crops University 570228.R.China;