Sichuan Zoige Alpine Wetland Ecosystem National Observation and Research Station Minzu University;
Institute of Qinghai-Tibetan Plateau Minzu University;
Plant Biology Section of Integrative Plant Science University;
Center for Forest Ecosystem Studies and Qianyanzhou Ecological Station Laboratory of Ecosystem Network Observation and Modeling of Geographic Sciences and Natural Resources Research Academy of Sciences;
Chengdu Institute of Biology Academy of Sciences;
State Key Laboratory of Grassland Agro-Ecosystem of Ecology University;
Nitrogen; Phosphorus; Scaling exponents; Twigs; Woody plants;