[1]State Key Laboratory of Protein and Plant Gene Research, College of Life Sciences, Center for Quantitative Biology, Peking University,Beijing 100871, China;
[2]Beijing International Center for Mathematical Research (BICMR) and Biomedical Pioneering Innovation Center (BIOPIC), Peking University,Beijing 100871, China;
[3]Department of Applied Mathematics, University of Washington,Seattle, WA 98195, USA;