The invention provides a plant in which, by virtue of modulation of the expression of a TETRASPORE (TES) or TES-like (TLK) gene, or of modulation of the activity of a protein encoded by such a gene, tetrad formation in the anther is disrupted, or is capable of being disrupted, such that callose cross-wall formation in the tetrad fails to the extent that fusion between two or more of the four microspore nuclei may occur.
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