The leaves ofGoldfussia dalhousianabecome infected byPuccinia polliniaewith the onset of rains in the Western Himalayas. Pale yellow dots appear on the leaves, developing into pustules and extending centrifugally. Pycnia develop on the upper surface and aecia on the lower. Protoaecia are differentiated into fertile and displacement zones. After dikaryotization, the basal cells elongate, become binucleate, and, by transverse conjugate division, produce the aeciospore mother cells. By transverse division, each of these form a large aeciospore and a small disjunctor cell towards the basal cell. The basal cells at the periphery, by transverse conjugate divisions, give peridial mother cells each of which by an oblique conjugate division cuts off an intercalary cell towards the outside and a peridial cell towards the inside. The peridial cells develop thick walls and the intercalary cells degenerate. The peridium has a dual origin and the aecia are of the aecidioid type. The aeciospores are circular to oval in shape, binucleate, and multiguttulate. They have radially striate thickened walls and two to three germ pores.
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