Semiaquatic bugs: phylogeny and classification of the Hebridae (Heteroptera: Gerromorpha) with revisions ofTimasius, NeotimasiusandHyrcanus

摘要

AbstractVarious characters of semiaquatic bugs belonging to the family Hebridae are analysed with respect to their taxonomic and phylogenetic importance. The cladistic relationships of the genera are discussed and a cladogram included. A classification is presented which divides the family into two sub‐families and resurrectsLipogomphusBerg as a valid taxon (to includeL. lacuniferusBerg (type‐species),L.accola(DrakeChapman)comb.n.forMerragata accola, L.brevis(Champion)comb.n.forMerragata brevis, andL.leucosticta(Champion)comb.n.forMerragata leucosticta). The following new taxa of hygropetric hebrids are described:Timasius livenssp.n.(Thailand),T.minorsp.n.(Pakistan, N. India),T.miyamotoisp.n.(Thailand),T.majorsp.n.(N.India),T.ventralissp.n.(N. India),T. miyamotoisp.n.(S. India),T.rupestrissp.n.(N. India),T.falcifersp.n.(N. India),T.championisp.n.(N. India),T.himalayensissp.n.(N. India),Neotimasius orientalisgen.n., sp.n.(S. India), Hyrcaninaesubfam.n.,Hyrcanus disparsp.n.(N. India),H.saxatilissp.n.(Thailand) andH.varicolorsp.n.(Java, Sumatra). Other species of the generaTimasiusandHyrcanusare redescribed and the generic position of one species changed:Hebrus(s.str.)atratus(Distant)comb.n.forTimasius atra