In a recent paper, MCiller and Reisz(1) proposed how fossil calibrations should be selected for application in molecular clock studies. The topic is of interest to those, like ourselves, who are actively engaged in reconstructing the tree of life using molecules. Nonetheless, we believe that these authors have erred both in their proposal of fossil calibration constraints and in their characterization of debates in the field. Their paper is an extension of one such debate that took place in another journal, where we responded to some of their initial criticisms' of our work. Here, we respond to their additional criticisms' and to their proposal for constraining fossil calibrations.
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