Revision of the European Rhyacophila fasciata species complex by fine phenomics of the paramere (Trichoptera, Rhyacophilidae).

摘要

The poorly known, so called widely distributed and highly varying species of the European Rhyacophila fasciataspecies complex are revised by fine phenomics of the paramere organisation. In this species complex paramere is the mostdiverse organ. It is the speciation trait integrating the initial split of speciation by its stimulatory and titillating functioninvolved in the early processes of reproductive isolation. Based on paramere organisation and on the character state ofdistribution three lineages have been delineated in the Rhyacophila fasciata species complex: the European R. fasciatalineage, the Caucasian R. aliena lineage and the R. mysica lineage distributed from Albania to Pakistan. In the R. fasciatalineage we have distinguished three clades of species: R. fasciata with five species, R. matrensis with seven species, and R.denticulata with seven species. We have re-diagnosed three known species: R. fasciata Hagen, 1859, R. denticulataMcLachlan, 1879, R. sociata Navas, 1916 and described 15 species new to science: R. biharensis Oláh sp. nov., R. bulgaricaOláh sp. nov., R. coppai Oláh sp. nov., R. csornahorensis Oláh & Szcz?sny sp. nov., R. ferda Oláh sp. nov., R. kopasa Oláh& Coppa sp. nov., R. matrensis Oláh, & Szcz?sny sp. nov., R. retezatensis Oláh sp. nov., R. rova Oláh & Coppa sp. nov. R.ruda Oláh & Johanson sp. nov., R. salfa Oláh sp. nov., R. soreda Coppa & Oláh sp. nov. R. suna Oláh sp. nov., R. tuhegaOláh sp. nov., R. zemplenensis Oláh sp. nov. The species status of Rhyacophila gemella Navas, 1923 was reinstated.