School of Forestry;
Northeast Forestry University;
Harbin 150040;
China;
College of Agricultural Resource and Environment;
Heilongjiang University;
Harbin 150080;
China;
College of Life Sciences;
Henan University;
Kaifeng 475004;
China;
institute of Applied Ecology;
Chinese Academy of Sciences;
Shenyang 110016;
China;
copiotrophic bacteria; denitrification; multidimensional effect; nitrification; oligotrophic bacteria; seasonal changes; soil acidification; water stress;