College of Fisheries Ocean University of China Qingdao 266003 China;
College of Fisheries Ocean University of China Qingdao 266003 ChinaField Observation and Research Station of Haizhou Bay Fishery Ecosystem Ministry of Education Qingdao 266003 China;
College of Fisheries Ocean University of China Qingdao 266003 ChinaField Observation and Research Station of Haizhou Bay Fishery Ecosystem Ministry of Education Qingdao 266003 China;
College of Fisheries Ocean University of China Qingdao 266003 ChinaField Observation and Research Station of Haizhou Bay Fishery Ecosystem Ministry of Education Qingdao 266003 China;
Marine College Shandong University Weihai 264209 China;
generalized additive model; principal component analysis; biotic variables; spatial autocovariate;