Key Laboratory of Mariculture(Ocean University of China);
Ministry of Education;
Agriculture Ministry Key Laboratory of Healthy Freshwater Aquaculture;
Key Laboratory of Freshwater Aquaculture Genetic and Breeding of Zhejiang Province;
Zhejiang Institute of Freshwater Fisheries;
L.maculatus; hCG; GnRHa; salinity; steroidogenic; enzyme; spermatogenesis;