Key Laboratory of Tropical Marine Bio-Resources and Ecology,South China Sea Institute of Oceanology,Chinese Academy of Sciences,Guangzhou 510301,China;
State Key Laboratory of Tropical Oceanography,South China Sea Institute of Oceanology,Chinese Academy of Sciences,Guangzhou 510301,China;
Key Laboratory of Marginal Sea Geology,Guangzhou Institute of Geochemistry,Chinese Academy of Sciences,Guangzhou 510640,China;
College of Animal Science and Technology,Jiangxi Agricultural University,Nanchang 330045,China;