Department;
of;
Bioengineering,;
University;
of;
Illinois;
at;
Chicago,;
Chicago,;
IL;
60607,;
U.S.A;
Shanghai;
Center;
for;
Systems;
Biomedicine,;
Shanghai;
Jiao;
Tong;
University,;
Shanghai;
200240,;
China;
Department;
of;
Applied;
Mathematics,;
University;
of;
Washington,;
Seattle,;
WA;
98195,;
U.S.A;
Kavli;
Institute;
for;
Theoretical;
Physics;
China,;
Chinese;
Academy;
of;
Sciences,;
Beijing;
100190,;
China;