National;
Key;
Laboratory;
of;
Plant;
Physiology;
and;
Biochemistry;
CAU-UCR;
Joint;
Center;
for;
Biological;
Sciences,;
China;
Agricultural;
University,;
Beijing;
100094,;
China;
Center;
for;
Plant;
Cell;
Biology;
and;
Department;
of;
Botany;
and;
Plant;
Sciences,;
University;
of;
California,;
Riverside,;
CA;
92521,;
USA;