Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu 610041, China;
Synthesis Research Centre of Chinese Ecosystem Research Network, Key Laboratory of Ecosystem Network Observation and Modelling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China;
Land and Resource College, China West Normal University, Nanchong 637002, China;
Institute of Qinghai-Tibetan Plateau, Southwest University for Nationalities Sichuan, Chengdu 610041, China;
Qinghai Academy of Animal Science and Veterinary Medicine, Xining 810016, China;