Jilin Engineering Laboratory for Avian Ecology and Conservation Genetics School of Life Sciences Northeast Normal University Changchun 130024 China;
Ministry of Education Key Laboratory of Vegetation Ecology School of Life Sciences Northeast Normal University Changchun 130024 China;
Jilin Provincial Key Laboratory of Animal Resource Conservation and Utilization School of Life Sciences Northeast Normal University Changchun 130024 China;
Ministry of Education Key Laboratory of Vegetation Ecology School of Life Sciences Northeast Normal University Changchun 130024 China;
Ministry of Education Key Laboratory for Ecology of Tropical Islands College of Life Sciences Hainan Normal University Haikou 571158 China;
Jilin Engineering Laboratory for Avian Ecology and Conservation Genetics School of Life Sciences Northeast Normal University Changchun 130024 China;
Ecologie Systématique Evolution Université Paris-Sud CNRS AgroParisTech Université Paris-Saclay Orsay Cedex F-91405 France;
Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering College of Life Sciences Beijing Normal University Beijing 100875 China;
