螅状独缩虫表膜下纤毛系及形态发生

摘要

利用蛋白银技术研究了螅状独缩虫无性生殖周期中的形态发生。结果表明: (1)在大核形态尚未出现明显变化时，生发毛基索(GK)的前端即出现原基, 随后原基增生扩大。最早出现的是两条将来分别演化为新仔虫第三咽膜(P′3)和第二咽膜(P′2)的原基带, 随后出现的是位于外侧的新仔虫的单毛基索(HK′)。同时，新仔虫的第一咽膜(P′1)也开始由老单毛基索(HK)复制，并在细胞分裂后期与老结构分离; (2)大核在虫体分裂过程中由长带状逐渐缩短变粗至扁圆形, 于虫体即将分开时迅速拉长，然后分裂为二个新大核; 小核分裂先于大核，在两仔虫口纤毛器即将分开时完成; (3) 原帚胚及柄归属老仔虫, 新仔虫的帚胚在虫体分裂后逐渐形成, 柄内肌丝则在柄鞘形成后逐渐长出。%Infraciliature and morphogenesis of Carchesium polypinum during asexual cucle were studied with protargol technique.The results show:(1)Before the obvioudty changed to the macronucleus in the front of the germinal row proliferating.With the development of the kinetosomes within the field,the peniculus 2 and 3 are formed,later the new haplokinety appeared,while the old haplokinety generated or duplicated the new peniculus 1.Both the new peniculus 1 and the old haplokinety separated at the telophase.The two germinal rows germinal rows(GK＇)for both the daughter cells appeared almost at the same time; (2) The macronucleus became shorter and thicker, finally oblate during the division of the cell. Then the macronucleus restored its original shape, band-like form. The micronucleus were divided into two before the macronucleus division; (3) The original scopula and stalk contributed to the old cell. The new cell formed scopula gradually after the cell division.