Key Laboratory of Experimental Marine Biology Institute of Oceanology Center for Ocean Mega-Science Chinese Academy of Sciences Qingdao 266071 China;
Laboratory for Marine Biology and Biotechnology Qingdao National Laboratory for Marine Science and Technology Qingdao 266237 China;
University of Chinese Academy of Sciences Beijing 100049 China;
Tianjin Weishan Road Middle School Tianjin 300222 China;