Key Laboratory of Marine Bio-Resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China;
Tropical Marine Biological Research Station in Hainan, Chinese Academy of Sciences, Sanya 572000, China;
Graduate University of Chinese Academy of Sciences, Beijing 100049, China;
Key Laboratory of Marine Bio-Resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China;
Tropical Marine Biological Research Station in Hainan, Chinese Academy of Sciences, Sanya 572000, China;
Yantai Institute of Coastal Zone Research, Chinese Academy of Sciences, Yantai 264003, China;
Key Laboratory of Marine Bio-Resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China;