Effect of stand density and humus removal on regeneration of Pinus densiflora seedlings in a mixed conifer and broad-leaved forest

摘要

We studied the impact of stand density and forest floor humus removal on regeneration of Pinus densiflora in a mixed conifer and broad-leaved forest on the Daimonji-Yama Mountain,Kyoto,Japan.Three levels of stand density were set,i.e.,a clear-cut site,a heavily thinned site and a lightly thinned site.In each density treatment,comparisons were carried out between humus removal (A 0 -free)and humus intact(A 0 -intact)subplots.We counted the emergence of sown pine seeds and measured survival and growth of pine seedlings over the next four years.In addition,light conditions and the recovery of understory were monitored.Results show that thinning intensity and humus removal promoted the establishment and growth of seedlings.One exception,however,was that seedling growth was minimal in the heavily thinned A 0 -intact subplots.This was due to a strong response of understory vegetation growth on the treatment combination,which hindered the pine seedling growth.Furthermore,we found that the canopy openness measured at 1.3-m height remained at least 35%for the heavily thinned site in 2008.The results suggest that the removal of the A 0 layer after heavy thinning is the most effective and labor-saving operation for P.densiflora regeneration.