State Key Laboratory of Genetic Resources and Evolution Kunming Institute of Zoology Chinese Academy of Sciences Kunming 650223 Yunan China;
College of Life Sciences Yunnan University Kunming 650091 Yunan China;
Kunming Natural History Museum of Zoology Kunming Institute of Zoology Chinese Academy of Sciences Kunming 650223 Yunan China;
Department of Ecology College of Life Sciences Henan Normal University Xinxiang 453007 Henan China;
Research Center for ecological management and protection of the Yellow River Basin Xinxiang 453007 Henan China;
State Key Laboratory of Genetic Resources and Evolution Kunming Institute of Zoology Chinese Academy of Sciences Kunming 650223 Yunan China;
College of Life Sciences Yunnan University Kunming 650091 Yunan China;
State Key Laboratory of Genetic Resources and Evolution Kunming Institute of Zoology Chinese Academy of Sciences Kunming 650223 Yunan China;