黔南紫云地区发育了全球罕见的晚石炭世后生动物格架礁.主要的造礁生物--Formitchevella,Ivanovia cf.manchurica、叶状藻和"未名造礁生物"与附礁生物一起各自构成了不同的造礁生物群落,群落内部的生物存在着空间竞争和共栖,而群落之间还有替代的关系.种内竞争使Formitchevella个体不断生长,且有时为避免与相邻个体的空间竞争而发生弯曲.在种间竞争方面,"未名造礁生物"以缠绕和接触的方式与Ivanovia cf.manchurica之间存在着直接主动的空间竞争;Formitchevella与Ivanovia cf.manchurica则保持着均势的相持平衡型竞争;叶状藻还可以以个体数量的优势在空间竞争上压倒了珊瑚.叶状藻与腕足类之间为一类特殊的共栖关系,二者互惠互利,在生存空间上存在一定的竞争.在扁平村珊瑚礁的形成过程中,Formitchevella群落与其下部群落的更替为建造晚石炭世大型珊瑚礁奠定了条件.%The Late Carboniferous metazoan framework reefs are unusually well developed in Ziyun region, southern Guizhou Province, China, even in a global perspective. The main reef-building organisms include Formitchevella, Ivanovia cf. manchurica, phylloid algae, and "unnamed reef-building organisms", and along with other inhabitants they constitute different reef communities. Within a given community, the organisms compete for space while maintaining symbiosis. In addition, ecological replacement also occurs among the different communities. Intraspecies competition drives Formitchevella skeleton to grow continuously, and sometimes the skeleton may be curled to avoid the spatial competition with the neighbors. As for interspecies competition, the "unnamed reef-building organisms" and Ivanovia cf. manchurica maintain an active and direct spatial competition through enwinding and contacting each other. Formitchevella and Ivanovia cf. manchurica keep a balanced competition and reach a competitive equilibrium, whereas phylloid algae surpass corals with their sheer number of countless individuals. The phylloid algae and the brachiopods have special ecological interactions, i. e. , maintaining symbiosis while still competing for living space. In the reef-building process of Bianping village coral reef, the replacement of Forrnitchevella community and its lower communities resulted in the formation of Late Carboniferous large-scale coral reef.
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