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Understory Species Patterns and Diversity in Old-Growth and Managed Northern Hardwood Forests

机译:老生长和管理北方硬木林的林下物种模式和多样性

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摘要

Forest management can significantly affect both the diversity and spatial patterning of understory vegetation. However, few studies have considered both diversity and spatial patterning at a stand scale. Our objective was to evaluate the effects of forest management on understory plant communities in northern hardwood forests and assess the processes governing differences in species composition, diversity, and spatial patterns. We sampled understory vegetation (all speciestall) and percentage of light transmission levels in three forest types in 12 mesic northern hardwood stands in northern Wisconsin and the Upper Peninsula of Michigan, USA: old-growth, undisturbed forests; even-aged forests resulting from clearcut logging (65–85 yr old); and uneven-aged forests with recent selective logging.Estimated understory species richness per stand, mean species richness per quadrat, and mean percent cover per quadrat were lower in old-growth forest than in even-aged, second-growth forests and lower in even-aged than in uneven-aged, second-growth forests. Differences in species composition among the three forest types were related to available light and to coarse woody debris; however, differences between the cover of most plant groups were not significant. The mean patch size of species diversity and cover is highly variable and could not be related to forest stand type. However, understory communities in old-growth forests have significantly smaller community patch sizes and larger compositional heterogeneity. Community patch size is correlated with both coarse woody debris and light heterogeneity. Each forest stand type produces a characteristic combination of understory composition, diversity, and spatial patterning of communities. Although harvesting has negligible effects on understory alpha diversity in these mesic hardwood forests, spatial structure is slower to recover and has not recovered in the even- and uneven-aged northern hardwood forests studied. If management objectives include preserving or restoring the ecological character of the forest, harvesting may need to be altered or delayed predicated on the character of the understory.
机译:森林管理可以显着影响林下植被的多样性和空间格局。但是,很少有研究考虑到林分规模的多样性和空间格局。我们的目标是评估森林管理对北部硬木林林下植物群落的影响,并评估控制物种组成,多样性和空间格局差异的过程。我们在美国威斯康星州北部和密歇根州上半岛的12种中等北部硬木林中,对3种森林类型的林下植被(所有种高)和透光率的百分比进行了采样:原始,未受干扰的森林;伐木(65-85岁)造成的平均年龄的森林;老龄林的估计林分平均林分丰富度,平均林分平均蓄积量和平均每平方英亩覆盖率均低于同龄,次生林,甚至更低。年龄比不均匀年龄的次生林中的年龄大。三种森林类型之间物种组成的差异与可用光和粗木屑有关。但是,大多数植物类别之间的覆盖率差异并不显着。物种多样性和覆盖度的平均斑块大小变化很大,并且与林分类型无关。但是,旧林中的林下群落具有明显较小的群落斑块大小和较大的成分异质性。群落斑块的大小与粗木屑和轻度异质性相关。每种林分类型都会产生林下组成,多样性和社区空间格局的特征组合。尽管采伐对这些内陆硬木林林下α多样性的影响可忽略不计,但在研究的均匀和不均年龄的北部硬木林中,空间结构的恢复较慢且尚未恢复。如果管理目标包括保护或恢复森林的生态特性,则可能需要根据林下的特性来改变或推迟采伐。

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