Males may use sexual displays to signal their quality to females; the handicap principle provides a mechanism that could enforce honesty in such cases. Iwasa et al. udmodel the signalling of inherited male quality, and distinguish between three variants of the handicap principle: pure epistasis, conditional, and revealing They argue that only the second and third will work. An evolutionary simulation is presented inudwhich all three variants function under certain conditions; the assumptions made by Iwasa et al. are questioned.
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