A phylogeny of the family Fanniidae Schnabl (Insecta : Diptera :Calyptratae) based on adult morphological characters, with specialreference to the Austral species of the genus Fannia

摘要

The present study proposed a phylogenetic hypothesis of the family Fanniidae based on a cladistic analysis usingcharacters from adult external morphology and female and male terminalia. The main purpose of this study was to clarify thephylogenetic position of newly described or poorly known species, mostly from southern South America, the Neotropics,Africa, Australia and New Zealand. In total, 151 characters from adult male and female external morphology and terminaliawere scored for 78 species of Fanniidae. Ten continuous characters were included and analysed as such. Three genera ofFanniidae and all the species-groups and subgroups proposed for the genus Fannia, except for the admirabilis-group and thesetifer-subgroup were included as terminal taxa. An heuristic parsimony analysis under implied weights was performed. Theanalysis recovered the monophyly of the Fanniidae and the genus Fannia, as well as the monophyly of several species-groupswithin Fannia. Male and female external morphological characters were, in general, highly homoplasious, whereascharacters from male terminalia showed low level of homoplasy and provided resolution at suprageneric nodes andspecies-groups.