Pterosaur track assemblages from the Upper Jurassic (lower Kimmeridgian) intertidal deposits of Poland: Linking ichnites to potential trackmakers

摘要

Two distinct tetrapod track assemblages, dominated by pterosaur traces, are reported from Upper Jurassic (lower Kimmeridgian, Hypselocyclum Zone) tidal flat carbonate deposits exposed in Wierzbica Quarry, 20 km south of Radom, Poland. The pterosaur tracks, tentatively assigned to Pteraichnus isp. are well preserved as a positive hyporelief in intertidal facies. Some are preserved with anatomical details (e.g., skin, claw or digital pad impressions) and quality sufficient to make preliminary ichnotaxonomic assignment. Nearly all collected specimens are larger than most other pterosaurian ichnites of the Late Jurassic age, thus adding to the growing diversity of pterosaur traces known from this time period. Morphometrical and anatomical analyses show that ctenochasmatoids or possibly non-pterydactyloid monofenestratans or rhamphorhynchids were most likely the trackmakers. Anatomical comparisons give new insights into the morphology of the pes of pterydactyloids and the most closely related non-pterodactyloids which may be correlated with a niche expansion into marginal marine and lacustrine environments by Middle and Late Jurassic pterosaurs. The palaeoenvironmental interpretation as tidal flat is confirmed by presence of vertically accreted tidal bundles, each representing deposition in one tidal cycle. Most likely then, this was one of the environments frequented by these pterosaurs and other small tetrapods. The wider palaeoecological significance of the tracks is evaluated in the context of other similar reports from the Late Jurassic of Europe, Africa and North America.