West African earthworm genus Millsonia Beddard, 1894 (Oligochaeta: Acanthodrilidae, Benhamiinae) reviewed and separation of a new genus

摘要

A detailed morphological revision of the species belonging to the genus Millsonia Beddard, 1894 is presented, and a new species Millsonia bouna sp. n. is described. To accommodate the species possessing enteronephric excretory system a new genus Reginaldia gen. nov. is erected. The new genus is divided into two subgenera according to the number of oesophageal gizzards: Reginaldia (Reginaldia) subgen. nov. for the species possessing two gizzards in v and vi, and Reginaldia (Peritogaster) subgen. nov. for the species provided with only one gizzard in vi.