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A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)

机译:Tribe Hygrotini Portevin的新分类,1929年(鞘翅目:Dytiscidae:Hydroporinae)

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摘要

The tribe Hygrotini Portevin, 1929 is currently composed of four genera, Heroceras Guignot, 1950, Herophydrus Sharp, 1880, Hygrotus Stephens, 1828 (with two subgenera, Coelambus Thomson, 1860, and Hygrotus s. str.), and Hyphoporus Sharp, 1880. A recent molecular phylogeny of the tribe with ca. 45% of the 137 described species of Hygrotini, including the type species of all genus-level taxa, revealed extended para-or polyphyly of the current genera and subgenera (Villastrigo et al., Zoologica Scripta, in press), for which reason a new classification of the tribe Hygrotini is proposed. Within Hygrotini only two genera are recognised: Clemnius n. gen. (with two subgenera: Clemnius s. str. with type species Hyphydrus decoratus Gyllenhal, 1810, and Cyclopius n. subgen. with type species Hydroporus acaroides LeConte, 1855) and Hygrotus (with four subgenera: Coelambus, Hygrotus s. str., Hyphoporus n. stat. and Leptolambus n. subgen. with type species Dytiscus impressopunctatus Schaller, 1783). Two genera are synonymised under Hygrotus s. str., Herophydrus n. syn. and Heroceras n. syn. The following 67 new combinations, for species thus far treated under the genera Heroceras, Herophydrus, Hygrotus and Hyphoporus, result from the new classification: Clemnius (s. str.) berneri (Young & Wolfe, 1984) n. comb., Clemnius (s. str.) decoratus (Gyllenhal, 1810) n. comb., Clemnius (s. str.) hydropicus (LeConte, 1852) n. comb., Clemnius (s. str.) laccophilinus (LeConte, 1878) n. comb., Clemnius (s. str.) sylvanus (Fall, 1917) n. comb., Clemnius (Cyclopius) acaroides (LeConte, 1855) n. comb., Clemnius (Cyclopius) farctus (LeConte, 1855) n. comb., Clemnius (Cyclopius) marginipennis (Blatchley, 1912) n. comb., Hygrotus (s. str.) assimilis (Regimbart, 1895) n. comb., H. (s. str.) bilardoi (Bistrom & Nilsson, 2002) n. comb., H. (s. str.) capensis (Regimbart, 1895) n. comb., H. (s. str.) confusus (Regimbart, 1895) n. comb., H. (s. str.) descarpentriesi (Peschet, 1923) n. comb., H. (s. str.) discrepatus (Guignot, 1954) n. comb., H. (s. str.) endroedyi (Bistrom & Nilsson, 2002) n. comb., H. (s. str.) gigantoides (Bistrom & Nilsson, 2002) n. comb., H. (s. str.) gigas (Regimbart, 1895) n. comb., H. (s. str.) goldschmidti (Pederzani & Rocchi, 2009) n. comb., H. (s. str.) gschwendtneri (Omer-Cooper, 1957) n. comb., H. (s. str.) hyphoporoides (Regimbart, 1895) n. comb., H. (s. str.) ignoratus (Gschwendtner, 1933) n. comb., H. (s. str.) inquinatus (Boheman, 1848) n. comb., H. (s. str.) janssensi (Guignot, 1952) n. comb., H. (s. str.) kalaharii (Gschwendtner, 1935) n. comb., H. (s. str.) morandi (Guignot, 1952) n. comb., H. (s. str.) muticus (Sharp, 1882) n. comb., H. (s. str.) natator (Bistrom & Nilsson, 2002) n. comb., H. (s. str.) nigrescens (Bistrom & Nilsson, 2002) n. comb., H. (s. str.) nodieri (Regimbart, 1895) n. comb., H. (s. str.) obscurus (Sharp, 1882) n. comb., H. (s. str.) obsoletus (Regimbart, 1895) n. comb., H. (s. str.) ovalis (Gschwendtner, 1932) n. comb., H. (s. str.) pallidus (Omer-Cooper, 1931) n. comb., H. (s. str.) pauliani (Guignot, 1950) n. comb., H. (s. str.) quadrilineatus (Regimbart, 1895) n. comb., H. (s. str.) reticulatus (Pederzani & Rocchi, 2009) n. comb., H. (s. str.) ritsemae (Regimbart, 1889) n. comb., H. (s. str.) rohani (Peschet, 1924) n. comb., H. (s. str.) rufus (Clark, 1863) n. comb., H. (s. str.) sjostedti (Regimbart, 1908) n. comb., H. (s. str.) spadiceus (Sharp, 1882) n. comb., H. (s. str.) sudanensis (Guignot, 1952) n. comb., H. (s. str.) travniceki (Stastny, 2012) n. comb., H. (s. str.) tribolus (Guignot, 1953) n. comb., H.
机译:部落Hygrotini波特万,1929目前由四个属,Heroceras Guignot,1950年,Herophydrus夏普,1880年,Hygrotus斯蒂芬斯,1828(有两个亚属,Coelambus汤姆森,1860年,和Hygrotus第STR),和Hyphoporus夏普,1880一种最近用约部落的分子系统发生Hygrotini的137个描述的物种的45%,包括所有属级分类单元的类型的物种,显示扩展的对位或多元性当前属和亚属(Villastrigo等人,动物SCRIPTA,出版中),为此原因部落Hygrotini新提出分类。在Hygrotini只有两个属被认可:Clemniusñ。根本。 (带有两个亚属:。Clemnius小号STR型物种Hyphydrus decoratus Gyllenhal,1810,和CyclopiusÑ亚属类型物种Hydroporus acaroides蠹,1855。)和Hygrotus(四个亚属:。Coelambus,Hygrotus小号STR,Hyphoporus ñ。STAT。和Leptolambusñ。用型物种Dytiscus impressopunctatus沙勒,1783)亚属。两个属正在Hygrotus小号synonymised。海峡,Herophydrusñ。 syn。和Herocerasñ。 syn。下列67个新的组合,种属Heroceras,Herophydrus,Hygrotus和Hyphoporus,从新的分类结果下迄今处理过:(:S STR)Clemnius berneri(杨&沃尔夫,1984)N。梳,Clemnius(一个或多个。STR。)decoratus(Gyllenhal,1810)N。梳,Clemnius(一个或多个。STR。)hydropicus(蠹,1852)N。梳,Clemnius(一个或多个。STR。)laccophilinus(蠹,1878)N。梳,Clemnius(一个或多个。STR。)西尔韦纳斯(秋季,1917)N。梳,Clemnius(Cyclopius)acaroides(蠹,1855)N。梳,Clemnius(Cyclopius)farctus(蠹,1855)N。梳,Clemnius(Cyclopius)marginipennis(Blatchley,1912)N。梳,Hygrotus(一个或多个。STR。)assimilis(Regimbart,1895)N。梳子。,H.(S。STR。)bilardoi(Bistrom&尼尔森,2002)N。梳子。,H.(S。STR。)兔(Regimbart,1895)N。梳子。,H.(S。STR。)confusus(Regimbart,1895)N。梳子。,H.(S。STR。)descarpentriesi(Peschet,1923)N。梳子。,H.(S。STR。)discrepatus(Guignot,1954)N。梳子。,H.(S。STR。)endroedyi(Bistrom&尼尔森,2002)N。梳子。,H.(S。STR。)gigantoides(Bistrom&尼尔森,2002)N。梳子。,H.(S。STR。)牡蛎(Regimbart,1895)N。梳子。,H.(S。STR。)goldschmidti(Pederzani&罗基,2009)N。梳子。,H.(S。STR。)gschwendtneri(奥马尔-库珀,1957)N。梳子。,H.(S。STR。)hyphoporoides(Regimbart,1895)N。梳子。,H.(S。STR。)ignoratus(Gschwendtner酒店,1933)N。梳子。,H.(S。STR。)inquinatus(Boheman,1848)N。梳子。,H.(S。STR。)janssensi(Guignot,1952)N。梳子。,H.(S。STR。)kalaharii(Gschwendtner酒店,1935)N。梳子。,H.(S。STR。)莫兰迪(Guignot,1952)N。梳子。,H.(S。STR。)孔雀(夏普,1882)N。梳子。,H.(S。STR。)natator(Bistrom&尼尔森,2002)N。梳子。,H.(S。STR。)变黑(Bistrom&尼尔森,2002)N。梳子。,H.(S。STR。)nodieri(Regimbart,1895)N。梳子。,H.(S。STR。)鲀(夏普,1882)N。梳子。,H.(S。STR。)obsoletus(Regimbart,1895)N。梳子。,H.(S。STR。)卵圆(Gschwendtner酒店,1932)N。梳子。,H.(S。STR。)苍白球(奥马尔-库珀,1931)N。梳子。,H.(S。STR。)pauliani(Guignot,1950)N。梳子。,H.(S。STR。)quadrilineatus(Regimbart,1895)N。梳子。,H.(S。STR。)网纹(Pederzani&罗基,2009)N。梳子。,H.(S。STR。)ritsemae(Regimbart,1889)N。梳子。,H.(S。STR。)鲁哈尼(Peschet,1924)N。梳子。,H.(S。STR。)鲁弗斯(克拉克,1863)N。梳子。,H.(S。STR。)sjostedti(Regimbart,1908)N。梳子。,H.(S。STR。)spadiceus(夏普,1882)N。梳子。,H.(S。STR。)sudanensis(Guignot,1952)N。梳子。,H.(S。STR。)travniceki(Stastny,2012)N。梳子。,H.(S。STR。)tribolus(Guignot,1953)N。梳子,H.

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