Taxonomic and nomenclatural notes on the Rhinusa tetra (Fabricius) species complex (Coleoptera: Curculionidae)

摘要

All taxa closely related to or synonymized with Rhinusa tetra (Fabricius, 1792) are studied, including the available type material. Four species are considered taxonomically valid: Rhinusa tetra, R. comosa (Rosenschoeld, 1838), R. moroderi (Reitter, 1906), R. verbasci (Rosenschoeld, 1838). The following four new synonymies are proposed: R. tetra (= Gymnetron eoum Rosenschoeld, 1838 syn. n.; = Cleopus uncinatus Dufour, 1843 syn. n.; = Cleopus verbasci Dufour, 1843 syn. n.); R. moroderi (= Gymnetron otini Hustache, 1946 syn. n). Neotypes are designated for Cionus amictus Germar, 1821, Cleopus uncinatus and Cleopus verbasci. Lectotypes are designated for Curculio teter, Gymnetron comosum, Gymnetron crassirostre Lucas, 1849, Gymnetron eoum, Gymnetron fuscescens Rosenschoeld, 1838, Gymnetron haemorrhoum Rosenhauer, 1847, Gymnetron moroderi, Gymnetron plagiellum Gyllenhal, 1838, Gymnetron trigonale Gyllenhal, 1838 and Gymnetron verbasci, all currently included in Rhinusa. A key separating the four validspecies is supported by diagnoses, biological notes, distributional data and illustrations. These new findings are important because R. tetra in the broad sense was proposed as a potential candidate for the biological control of invasive Common Mullein (Verbascum thapsus L.) in North America.